Due to rising sea levels and a rapidly declining human population, androids have been engineered to perform most menial tasks, allowing humans to focus on other things. Aside from providing manual labor and customer service, androids also entertain humans through Babel—a tower where participants captivate audiences with song and dance to clear Climb Stages. The higher the climb, the more prize money is awarded. Androids Cobalt, Chrom, Neon, and Kei are down on their luck and about to have their electricity cut off from unpaid bills. Desperate to make a quick buck, the group decide to become Babel climbers. However, winning the audience's favor is a great challenge. Dejected but unwilling to give up, Cobalt resolves to somehow improve their chances after watching an impressive performance by STAND-ALONE—the top climber unit. When the group stumbles across a mysterious choreographer willing to give away his routine, Cobalt sees this as their only way up. With the help of their human companion, Esora Shibaura, the androids form the unit KNoCC to tackle the daunting climb to the top of Babel, but they must also overcome android-based discrimination along the way.

Arisugawa Academy's DJ unit Lyrical Lily is known for bringing smiles to people's faces through music. Thanks to their efforts, the local city hall requests them to perform monthly for a year as part of Operation Beckoning Cat, the town revitalization project. Unfortunately, given the long and demanding event, Lyrical Lily wonders if they can successfully meet the expectations placed on them. To better their chances, they decide to invite the DJ units that supported them during their first performance: Happy Around, Photo Maiden, Peaky P-key, Rondo, and Merm4id. Intrigued by the opportune event, the other units gladly accept Lyrical Lily's request. However, the project is extremely demanding. To attract visitors and new residents to the town, the energetic girls of the participating DJ units must give their best to create a memorable experience for their audiences.

Akira Kiyose, a high schooler who loves to sing, gets recruited by a talent agency called sMiLea Production. Akira learns that the legendary idols "Anela," who shocked the world with their sudden retirement, are behind the agency developing talent. Together with Banri Naoe and Chihiro Isuzugawa, also recruited by the agency, they form an idol group as they chase their dream towards super stardom alongside other idol groups "LEGIT" and "JAXX/JAXX."

A special episode focusing on the story of Merm4id and Rondo, including a new, joint performance by the two groups.