11-year-old Persia has grown up alongside the animals on the Serengeti plains of Africa. Twins Riki and Gaku Muroi and their grandfather, Gouken, bring Persia to Japan with them, where she will live with a couple (who Persia calls Mama and Papa) that own a grocery store. During the flight to Japan, Persia finds herself in the "Lovely Dream" where she meets the Fairy Queen, who explains that the Lovely Dream is in danger and requests Persia's help. Giving Persia a magical headband (which can also summon a magical baton) that allows her to transform into a teenage girl, connecting her to the Lovely Dream and allowing her to use magic. She is sent with three kappa back into the regular world with the mission of collecting love energy to thaw the frozen Lovely Dream. If Persia fails her task of collecting love energy or if someone catches her using magic, Riki and Gaku will be transformed into women.

Before Chise became the "ultimate weapon," there was another—Lieutenant Mizuki. A battle-hardened military woman, she volunteered for an experimental procedure after injuries left her unable to return to the battlefield. As the prototype ultimate weapon, Lieutenant Mizuki was highly successful on the battlefield, however, as the first candidate, her development was limited. When a more suitable candidate to become the weapon, Chise, is forced into the military, Lieutenant Mizuki thinks that she is silly, weak, and unsuited for the role. As the only other person to have undergone the procedure, however, Lieutenant Mizuki can hear Chise's thoughts and is the only one who understands her. As the war rages on and Chise's development progresses, Lieutenant Mizuki discovers more about Chise, ultimate weapons, and herself.

After his father's sudden death, Ceddie moves to England as an heir to his grandfather, the Earl of Dorincourt. The Earl is a stubborn and selfish old man. Gradually, however, Ceddie's innocent love opens the Earl's heart and changes him into a caring and generous person. This is based on "Little Lord Fauntleroy" by F.H.Burnett.

Second season to the original Tiger Mask, with a different cast of characters.