It adapts the story from volume 12 (chapter 98 onwards) of the manga series "Demon Slayer: Kimetsu no Yaiba" . The story takes place in a new setting - Tanjiro arrives at the "Swordsmith Village." After the death of Upper Moon Six and Daki, a vacancy appears among the upper moons, and Muzan Kibutsuji summons the remaining upper moons to the Infinity Castle. Muzan is furious about the vacancy and believes that the upper moons should do everything in their power to find the "Blue Spider Lily" and eliminate the Demon Slayer Corps. As a result, Upper Moon Four, Hantengu, and Upper Moon Five, Gyokko, are dispatched to carry out the mission. Meanwhile, Tanjiro, who has been recovering from his injuries at the Butterfly Estate for two months, learns that Kozo Kanamori, the swordsmith, has refused to make him a new sword. Following the advice of the Hashira, Shinobu Kocho, Suma Nakahara, and Kiyo Terauchi, Tanjiro travels to the Swordsmith Village to conUpon arrival, he encounters the Love Hashira, Mitsuri Kanroji, and the Mist Hashira, Muichiro Tokito, as well as Genya Shinazugawa. Tanjiro and his companions are about to embark on a new battle. The opening theme song "Ketsui no Hikari" is performed by MAN WITH A MISSION and milet. The lyrics and composition are by Jean-Ken Johnny, with arrangement by MAN WITH A MISSION and cooperation by Chisato Ichinose. The ending theme song "Koikogare" is also performed by MAN WITH A MISSION and milet.

Unaired episode included with the Blu-ray.

In the near future, humanity has taken its first steps towards journeying into the far reaches of the galaxy. Upon doing so they discover a huge race of insectoid aliens known as "Space Monsters." These aliens seem dedicated to the eradication of mankind as they near closer and closer to discovering Earth. In response, humanity develops giant fighting robots piloted by hand-picked youth from around the world. Shortly after the discovery of the aliens, Noriko Takaya, the daughter of a famous deceased space captain, enters a training school despite her questionable talents as a pilot. There, she meets her polar opposite, the beautiful and talented Kazumi Amano, and is unexpectedly made to work together with her as they attempt to overcome the trauma of war as well as their own emotions.

Upon first seeing Earth from the Azatos Empire's survey ship, Chief Investigator Liza Luna plans to eradicate every lifeform on it immediately, leaving behind only useful resources. While surveying the planet, Liza discovers that human civilization has evolved at a pathetic rate compared to other lifeforms. Liza makes her way to the surface of the planet to properly inspect and understand its inhabitants. Still convinced that there is nothing beneficial to be found, the investigator makes her way into a cafe to gauge the planet's food culture. Liza walks into a cat cafe, where she meets creatures far cuter than anything else in the galaxy. Motivated by this new discovery, Liza decides to investigate Earth's numerous adorable inhabitants.