In their last year of middle school, the five band members of CRYCHIC went their separate ways after an unpleasant disbandment. Each wrestling with their own regrets, the girls enrolled in different high schools, hoping never to hear from one another again. Meanwhile, Anon Chihaya transfers to Haneoka Girls' Academy in the middle of the school year. As most students play in a band as a hobby, Anon desires to fit in and spends her first days trying to recruit members for her own band, envisioning herself as the group's vocalist. She befriends Tomori Takamatsu, a shy girl who seems interested but hesitant about joining. As it turns out, Tomori used to be part of CRYCHIC, an experience that has scarred her deeply. Through Tomori, Anon meets Taki Shiina and Soyo Nagasaki, both former members of the same band. Although Taki detests the idea of playing with Anon, Soyo is delighted to help her out. Now with Soyo by her side, Anon's dream of forming a band is within reach—if only she could find a few more members.

When a genius patissier dies in a tragic accident before achieving his dream of creating the world's greatest sweets, he is reincarnated as nine-year-old Pastry Mille "Pas" Morteln. Now in a medieval world brimming with swords and magic, Pas enjoys a peaceful life in the Fief of Morteln under the authority of his father, the military hero Lord Casserole. However, scarce ingredients render sweets a luxury, so Pas can only dream of baking. Once news arrives of approaching bandits, Pas must learn magic and fight to defend his land, all to one day create a confectionery paradise!

The series stars Hat, the descendant of a hero who fought an evil king and sealed Devildom. Hat falls into Devildom and finds he has to defeat the evil king.

The series will focus on Gamera facing off against five different Kaiju.