The fifth season of .

Now in his second year of high school, Rei Kiriyama continues pushing through his struggles in the professional shogi world as well as his personal life. Surrounded by vibrant personalities at the shogi hall, the school club, and in the local community, his solitary shell slowly begins to crack. Among them are the three Kawamoto sisters—Akari, Hinata, and Momo—who forge an affectionate and familial bond with Rei. Through these ties, he realizes that everyone is burdened by their own emotional hardships and begins learning how to rely on others while supporting them in return. Nonetheless, the life of a professional is not easy. Between tournaments, championships, and title matches, the pressure mounts as Rei advances through the ranks and encounters incredibly skilled opponents. As he manages his relationships with those who have grown close to him, the shogi player continues to search for the reason he plays the game that defines his career. [Written by MAL Rewrite]

Humanity is finally confronted with the threat of the Custodians—the ancient, sentient species held responsible for cursing humanity to speak different languages thousands of years ago. The Symphogear wielders—Hibiki Tachibana, Tsubasa Kazanari, Chris Yukine, Maria Cadenzavna Eve, Kirika Akatsuki, and Shirabe Tsukuyomi—are sent to the Antarctic in order to retrieve an ancient relic. After securing it and rescuing the scientific staff present there from a Coffin, the automated defense mechanism protecting it, the relic is given to American researchers due to international agreements. The criminal organization Noble Red, a remnant of the previously fought Bavarian Illuminati, starts targeting the relic. Will the Symphogear wielders and their supporting organization S.O.N.G. be able to foil the plans of the organizations conspiring against them? [Written by MAL Rewrite]