To escape the explosion of their ship caused by a terrorist attack, the parents of a young baby put him inside an escape pod and send him out before dying. The pod ends up on a planet inhabited by beings able to transform themselves into animals by using a special ring. The young baby is rescued by the royal family, and grows up to become the prince. When the planet is attacked by pirates, he will be brought by force to Earth and discover everything about his own origins, and forgotten brother.

Yuu, Persia, Mai and Yumi are together in a science fiction-like adventure. An awful monster assault young and beautiful girls, leaving them transformed into ugly hags: only our four magical girls with their powers can fly to the space and fight for Earth 's sake, so Yuu transforms into Creamy Mami, Mai into Magical Emi, Persia into Fairy and little Yumi draws space suits for all of them.

Hagiwara Mio is a 14-year-old TV actress who has gotten popular. One day on a set, she meets Kumagai Ichiya, who tells her that her acting "stinks." Mio is hurt by his comments, but she finds that she can't get Ichiya out of her mind. It turns out that Ichiya is a promising director, who directed a music video for Mio's friend Sawaki Aya, an idol singer. After the success of this music video, Ichiya is asked to direct a movie, and he wants Mio to be the heroine, because he feels that she is a "handsome" girl like the actresses of old. How will Mio cope with her feelings toward Ichiya?

Short flash animation series of Carole & Tuesday released on the official YouTube and Twitter channels.