A-ko and B-ko are bounty hunters in an untamed world. When a young heiress, C-ko, falls out of the sky and into their hair, they find themselves battling space pirates and galactic police. Turns out the high-pitched, hungry C-ko holds a secret—the spirit of an ancient evil—and is the center of a plot to destroy the universe!

Maeda Toshiie and Matsu visit Chousokabe Motochika, who governs Shikoku. They tell Motochika unreasonable demands from Nobunaga. Motochika rejects them and urges Maeda to betray Nobunaga. Meanwhile Keiji tries to convince Mouri Motonari to join the fight against Nobunaga, and is imprisoned as a result. Motochika and Motonari move against each other.

The stage sets in a world where guinea pigs became cars. The car that makes you relax, "Molcar". Having big round eyes, curvy butt and speedy limbs, they always explore the world with their innocent faces. Even during traffic jams, you can be eased just by looking at the butt of that Molcar in front of you. Even when they cause trouble, their furry cuteness will somehow make you forgive them!? Surrounded by situations only met by cars, it's a Mol-animation filled with cure, friendship, adventure and furry actions!

The Galactic Empire and the Free Planets Alliance have been locked in a seemingly endless war for more than a century and a half. In the Empire, a young Siegfried Kircheis meets Reinhard von Müsel and his older sister Annerose. Kircheis enjoys a happy friendship with the two beautiful blonde-haired siblings until the day that their alcoholic father, a lesser nobleman with low standing, accepts a request for Annerose to be sold as a concubine to the Kaiser. Although enraged, Reinhard is powerless to stop the whims of the Imperial Court. He and his father soon move away, leaving Kircheis behind.

A few years later, Reinhard returns in a military uniform and declares his intent to rise through the ranks and free his sister from sexual servitude. Kircheis joins Reinhard on this daring journey to save Annerose, yearning to forever stay by the side of his friends.

[Written by MAL Rewrite]