Clad in desert pink and the size of a mere child, the infamous "Pink Devil" mercilessly hunts down other players in the firearm-centered world of the virtual reality game Gun Gale Online. But in real life, this feared player killer is not quite who anyone would expect. A shy university student in Tokyo, Karen Kohiruimaki stands in stark contrast to her in-game avatar—in fact, she happens to stand above everyone else too, much to her dismay. Towering above all the people around her, Karen's insecurities over her height reach the point where she turns to the virtual world for an escape. Starting game after game in hopes of manifesting as a cute, short character, she finally obtains her ideal self in the world of Gun Gale Online. Overjoyed by her new persona, she pours her time into the game as LLENN, garnering her reputation as the legendary player killer. However, when one of LLENN's targets gets the best of her, she ends up meeting Pitohui, a skilled yet eccentric woman. Quickly becoming friends with Karen, Pitohui insists that LLENN participates in Squad Jam, a battle royale that pits teams against one another, fighting until only one remains. Thrust into the heated competition, LLENN must fight with all her wit and will if she hopes to shoot her way to the top.

Saenome are tasked with sealing spirits that possess objects and pose a threat to society. Known as tsukumogami, these spirits are generally non-violent, and saenome normally seal them after a simple verbal exchange. Traumatized by a previous deadly encounter with a malevolent spirit, Hyouma Kunato refuses to use peaceful methods and gets into physical confrontations with all tsukumogami that cross paths with him. Deeming this behavior unacceptable for a future clan leader, Hyouma's grandfather, Zouhei, proposes that he starts living with Botan Nagatsuki, a young woman whose family is composed of six tsukumogami—lest he be excommunicated. Determined to track down the spirit that took everything from him, Hyouma reluctantly accepts Zouhei's proposal. As unknown groups of evil spirits begin to jeopardize the balance between worlds, the young man must gain the trust of Botan and overcome his hatred for her mysterious family.

Kouta has girl troubles of the supernatural sort. For some reason, he keeps attracting the attention (and affections) of animal spirits! Having spent most of his life in the country, Kouta is understandably nervous when he moves in with his grandma to attend a high school in the big city. He hoped to make a good impression, but having Chizuru, a beautiful fox spirit, hanging off his arm didn't seem to be the sort of image he wanted to have. She's not alone in her love for Kouta, either. Nozomu, a wolf spirit, as well as other youkai have their sights set on the hapless country boy.